Some episodes have featured adaptions of well-known folktales or ancient myths or fables, while others have featured popular storybooks such as Click, Clack, Moo: Cows that Type, or shown the lions learning or benefiting from the lessons presented by the story.

The program's format is intended to promote literacy and reading in each episode, the lions introduce a picture book to the audience and read it.

Busterfield III Memorial Public Library” starring alongside characters such as Click, an electronic, anthropomorphic computer mouse, the Information Hen, who answers library calls, and Heath, a dinosaur who serves as the library’s thesaurus. The series focuses on a family of clumsy anthropomorphic lions operating and living in a large, busy library called, “The Barnaby B. The show ran from Apto November 22, 2010, taking the schedule slot held by The Puzzle Place upon the latter's debut. It is created by alumni of Sesame Street and several season 2 episodes, notably in the Dance in Smarty Pants music videos, had a few characters from Sesame Street guest appearing. The show won seven Daytime Emmy awards between 20. The show was a co-production between WGBH in Boston and Sirius Thinking, Ltd., in New York City, in association with Mississippi Public Broadcasting, the distributor from seasons 5–10, in Mississippi. Between the Lions is an American animated/live-action/puppet children's television series designed to promote reading.

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